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Gymnosperm Stem: Medullary Rays Three Year Pinus

cross section: Pinus stem magnification: 100x Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. cross section: Pinus stem magnification: 100x Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem.

Gymnosperm Stem: Medullary Rays in Three Year Pinus

cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Three Year Pinus

cross section: Pinus stem magnification: 40x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 40x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Phloem in Three Year Pinus

cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Three Year Pinus

cross section: Pinus stem magnification: 100x Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. cross section: Pinus stem magnification: 100x Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem.

Gymnosperm Stem: Pith in Two Year Pinus

cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Pith in Three Year Pinus

cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Cambium in Three Year Pinus

cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Two Year Pinus

cross section: Pinus stem magnification: 40x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 40x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Herbaceous Dicot Stem: Parenchymatous Pith in Older Helianthus

cross section: Older stem: Helianthus common name: Sunflower magnification: 100x All vascular tissues in Helianthus are the product of primary growth. There are no signs of secondary growth (growth rings or lateral meristems) as might be seen in woody stems. The uniseriate and cutinized epidermis contains large multicellular trichomes and occasional stomata. The cortex consists of an outermost hypodermis of heavy walled collenchyma, a deeper layer of parenchyma and a deepest band of endodermis (starch sheath). Within the stele the vascular bundles are arranged in a ring and separated from each other by wide medullary rays of parenchyma cells. The collaterally arranged vascular bundles are almost entirely primary phloem and xylem. Each bundle consists of a large outer supportive cap of sclerenchyma fibers (phloem fiber cap), a deeper layer of primary phloem with well-defined sieve tubes and companion cells, and a deepest layer of primary xylem. In between the xylem and phloem, a narrow band of cambium may be seen. In some preparations, the highly lignified cells walls of xylem and mature sclerenchyma are stained red orange. These cells are dead at maturity and can also be distinguished by a heavy cell wall and absence of cytoplasm. The center of the stem is occupied by a pith of parenchyma cells that contain numerous starch storing amyloplasts. cross section: Older stem: Helianthus common name: Sunflower magnification: 100x All vascular tissues in Helianthus are the product of primary growth. There are no signs of secondary growth (growth rings or lateral meristems) as might be seen in woody stems. The uniseriate and cutinized epidermis contains large multicellular trichomes and occasional stomata. The cortex consists of an outermost hypodermis of heavy walled collenchyma, a deeper layer of parenchyma and a deepest band of endodermis (starch sheath). Within the stele the vascular bundles are arranged in a ring and separated from each other by wide medullary rays of parenchyma cells. The collaterally arranged vascular bundles are almost entirely primary phloem and xylem. Each bundle consists of a large outer supportive cap of sclerenchyma fibers (phloem fiber cap), a deeper layer of primary phloem with well-defined sieve tubes and companion cells, and a deepest layer of primary xylem. In between the xylem and phloem, a narrow band of cambium may be seen. In some preparations, the highly lignified cells walls of xylem and mature sclerenchyma are stained red orange. These cells are dead at maturity and can also be distinguished by a heavy cell wall and absence of cytoplasm. The center of the stem is occupied by a pith of parenchyma cells that contain numerous starch storing amyloplasts.

Gymnosperm Stem: Annual Rings in Three Year Pinus

cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Cortical Resin Duct in Three Year Pinus

cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Three Year Pinus

cross section: Pinus stem magnification: 40x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 40x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Cork and Cortex in Three Year Pinus

cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Cambium in Two Year Pinus

cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Epidermis and Cortex in Two Year Pinus

cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous.

Gymnosperm Stem: Cork in Three Year Pinus

cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. cross section: Pinus stem magnification: 400x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins.

Gymnosperm Stem: Two Year Pinus

cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts. cross section: Pinus stem magnification: 100x During the first year of growth the cutinized epidermis is replaced by protective growth of cork rich periderm. The outer periderm consists of layers of cork cells, the phellem, which produces waterproofing suberin. Cork cells are dead at maturity. Deep to the phellem is a living layer of cork cambium or phellogen and beneath that, layers of cork parenchyma or phelloderm. Many cells in the periderm contain dark staining tannins. The cortex is divided into a thin outer hypodermis of lignified sclerenchyma cells and thicker inner cortex of thin walled parenchyma cells containing chloroplasts. Large resin ducts are surrounded by secretory parenchyma that produces resins and turpentines. Some cells enlarge into dark staining tyloses. Both endoderm and pericycle are inconspicuous. The vascular cylinder or stele in young stems consists of a ring of vascular bundles interspaced with medullary rays of parenchyma cells. Seasonal activity of the cambium replaces the isolated vascular bundles with well-defined annual rings of secondary phloem and xylem. Xylem is endarch with protoxylem found towards center of the stem and younger metaxylem towards the periphery of the stem. Protoxylem consists of annular and spiral tracheids with only tracheids found in metaxylem. True xylem vessels are lacking. Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. Phloem is endarch but annual growth the of stem makes it difficult to distinguish between older protophloem to the periphery and younger metapholem towards center of the stem. Phloem lacks companion cells, consisting entirely of sieve tubes and phloem parenchyma. Medullary rays in the secondary phloem include protein rich albuminous cells. A well-defined pith of parenchyma cells is interrupted by a few large resin ducts.

Gymnosperm Stem: Resin Duct in Three Year Pinus

cross section: Pinus stem magnification: 100x Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem. cross section: Pinus stem magnification: 100x Because of the greater production of xylem, the vascular cylinder is dominated by radially arranged rays of secondary xylem interspaced with medullary rays of parenchyma cells. Conspicuous resin ducts are present throughout the xylem.

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